ExYSTEMS LAB
At ExYSTEMS, we investigate how plants persist, adapt and maintain diversity under environmental extremes and global change. We combine macroecological and mechanistic approaches to understand the processes shaping biodiversity and the evolutionary resilience of plant communities, with a particular interest in extreme ecosystems such as cliffs, rocky habitats and alpine environments.
At broad scales, we explore the functional, (phylo)genetic and environmental drivers of multidimensional biodiversity, species distributions and phenotypic variation across environmental gradients and extremes. At the mechanistic level, we investigate how plants respond to harsh and changing conditions through phenotypic plasticity, transgenerational responses and functional strategies, and how these responses are mediated by biotic interactions, including plant–microbe interactions.
By integrating evolutionary ecology, functional ecology, microbial ecology and conservation biology, ExYSTEMS connects fundamental research with real-world conservation challenges. Ultimately, we aim to understand the processes that determine the vulnerability and resilience of biodiversity to global change, and to translate this knowledge into effective management and conservation actions for ecosystems harbouring unique and threatened biodiversity.
Research Associates
Previous Group Members
Mireia S. Díaz Amate
Research Technician (June-September 2025)
BSc and MSc STUDENTS
Claudia Alfaro (BSc thesis – February-July 2025). “Evolutionary patterns and intra- and interspecific differences in the germination of Petrocoptis A. Braun ex Endl.” (University Rey Juan Carlos).
Patricia García Inarejos (MSc thesis – January-September 2025). “Functional trait responses of Arbutus unedo L. to global change drivers revealed through historical herbarium records” (Universidad Autónoma de Madrid).
Sara Peinado (BSc thesis – March-November 2025). “Effect of climbing cessation on rupicolous flora (Zorreras Bajas, Loja, Granada)” (Universidad de Granada).
Angie Paola Carvajal (MSc thesis – January-December 2025). “Phenotypic variation and functional expression in endemic and threatened species of the genus Petrocoptis A. Braun ex Endl.: implications for the conservation of cliff-dwelling plants” (University Rey Juan Carlos).
Andrea Porru (BSc thesis – June 2025-April 2026). “Comparative analysis of cliff-dwelling plant communities in two national parks of the Iberian Peninsula” (University of Cagliary, Italy).
Loïc Bondu (Intership BSc student – May-July 2026). (University of Rennes, France).
Raquel de la Torre (BSc thesis – March-July 2026). “Phenotypic plasticity in response to environmental stress in cliff-specialist plants: the role of population elevation of origin in Petrocoptis” (University Rey Juan Carlos).
Alba M. Milán (BSc thesis – March-September 2026). “Reproductive strategy as a modulator of phenotypic plasticity in response to drought and rising temperatures in cliff specialist species of the genus Petrocoptis” (University Rey Juan Carlos).








